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To reveal the ERK activity dynamics in the intestinal epithelium, we used transgenic mice ubiquitously expressing a highly sensitive Förster resonance energy transfer (FRET) biosensor for ERK activity (EKAREV-NLS) (Fig. 1a)30. The small intestine of EKAREV-NLS mice was observed under an inverted two-photon excitation microscope (Fig. 1b).
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