We document the establishment of the Aegea–Anatolia/Eurasia plate boundary in Pliocene–Pleistocene time. Before 2 Ma, no localized plate boundary existed north of the Aegean portion of the Anatolia plate and the shear produced by the motion of Anatolia–Aegea with respect to Eurasia was distributed over the whole width of the Aegean – West Anatolian western portion. In 4.5 Ma, a shear zone comparable to the Gulf of Corinth was formed in the present Sea of Marmara.