Our model is based on the notion that membrane shape feeds back with cortical protein recruitment19,21,29,30. Such feedback can generate wave behavior30, and similar ideas have been proposed to account for traveling waves in other systems29,31. Here we set out to explore how such concept plays out in our own system. We aimed to construct a minimal model—including the most essential components as demonstrated by existing experiments4—to recapitulate the basic phenomena of our traveling waves (Fig.