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1 Introduction Aberrant DNA hypermethylation at promoter CpG islands (CGIs) silences tumor suppressor genes (TSGs) and drives malignant progression of human cancers [1-4]. These DNA methylation abnormalities are mitotically inherited through DNA methyltransferase 1 (DNMT1), whose recruitment to nascent DNA depends on the cofactor protein UHRF1 (ubiquitin like with PHD and RING finger domains 1) [5-8].
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