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1 Introduction A crucial feature of adaptive immunity is the ability of B cells to recognise and respond to invading pathogens with their membrane-bound antibody, the B-cell antigen receptor (BCR). T cell-dependent responses lead to the activation of B cells, and the formation of germinal centre (GC) structures where B cells undergo clonal expansion, and their BCRs undergo somatic hypermutation (SHM) and class switch recombination (CSR) [1].
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