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1 Introduction Biological invasions have emerged as a primary driver of biodiversity loss and economic damage (Mori et al. 2026). The impacts of alien species on native biota and human societies are escalating rapidly (Verma et al. 2024). A subset of naturalized species that has overcome dispersal barriers and established self-sustaining populations far from their original introduction sites is classified as invasive alien plants (Richardson et al. 2001).
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