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1 Introduction Vascular epiphytes contribute about 10% to the global vascular plant species richness (Zotz et al. 2021) and up to 50% in some tropical cloud forests (Kelly et al. 2004). Due to the varying light and moisture conditions in the forest canopy, specific adaptations are associated with epiphytic growth; for instance, velamen radicum (sponge-like root tissue), atmospheric water uptake, CAM photosynthesis and a specialised root microbiome (Zotz 2016).
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