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1 Introduction Biological macromolecules can adopt topologically complex structures that influence their stability and function. Among these, molecular knots represent one of the most striking topologies accessible to polymer chains. Knots arise spontaneously in long polymers such as double-stranded DNA (dsDNA) through thermal fluctuations [1], as well as from enzymatic processes in viruses [2], bacteria [3], and eukaryotes [4, 5].
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