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1 Introduction Tropical biodiversity is besieged by many threats, including the over-exploitation of forests (Maxwell et al., 2016), hunting (Tilker et al., 2019), pollution (Hölker et al., 2010), fire (Kelly et al., 2020), climate change (Sintayehu, 2018), invasive species (Doherty et al., 2016), and habitat destruction (Hanski, 2011).
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