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1 INTRODUCTION Smk1 is a mitogen activated protein kinase (MAPK) family member in Saccharomyces cerevisiae that controls the postmeiotic program of spore formation (Krisak et al., 1994). It is produced specifically during meiosis, as cells enter the first meiotic division (MI) and activated upon completion of the second meiotic division (MII) (Whinston et al., 2013).
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