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Abstract Among the evolutionary innovations of many succulents is a photosynthetic lifestyle, where stomatal gas exchange is decoupled from light-dependent carbon fixation. Many Crassulaceae leaf succulents form a stomatal morphotype consisting of kidney-shaped guard cells surrounded by three anisocytic subsidiary cells (SCs), whose function and development remained unknown. Here, we established Kalanchoë laxiflora as a developmental model.
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