Introduction The plant cuticle is a hydrophobic barrier covering most aerial surfaces in land plants, protecting them from desiccation, pathogen attack and tissue fusion (González Valenzuela et al., 2023). Cuticles consist of a cutin matrix embedded with polysaccharides and cuticular waxes and overlaid with epicuticular waxes typically made from very long chain fatty acids (VLCFAs) and their derivatives (Yeats & Rose, 2013). However, within this framework, cuticles show remarkable differences.