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1 INTRODUCTION Grasslands cover more than one third of the Earth's terrestrial habitats and 35% of agriculturally used land in Europe (Gibson & Newman, 2019; Smit et al., 2008). They provide essential goods and ecosystem services (Gibson & Newman, 2019) and include the biomes with the highest vascular plant-species richness at <50 m2 scale (Boch et al., 2020; Wilson et al., 2012).
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