1,2,3,4 1Program in Computational Biology and Bioinformatics, Yale University, New Haven, Connecticut 06520, USA; 2Department of Molecular Biophysics and Biochemistry, Yale University, New Haven, Connecticut 06520, USA; 3Department of Computer Science, Yale University, New Haven, Connecticut 06520, USA; 4Department of Statistics and Data Science, Yale University, New Haven, Connecticut 06520, USA Corresponding author: mark{at}gersteinlab.org Footnotes [Supplemental material is available for...