This is an early access version, the complete PDF, HTML, and XML versions will be available soon. Open AccessArticle by Natalie Coe 1,*, Obadiah Mulder 2 and Kenneth Mulder 3 1 Biology Department, University of North Carolina, Wilmington, NC 28403, USA 2 Quantitative and Computational Biology, University of Southern California, Los Angeles, CA 90007, USA 3 Mathematics and Statistics, Mount Holyoke College, South Hadley, MA 01075, USA * Author to whom correspondence should be addressed.