1 Introduction There is no doubt that mesoscale eddies are prevalent nearly everywhere in the world’s oceans (Chelton et al., 2011; Chelton et al., 2007). Biogeochemical variables and processes are strongly influenced by mesoscale eddies (McGillicuddy et al., 1999; Patel et al., 2020). There is generally an increase in primary production in cyclonic cold eddies and a decrease in primary production in anticyclonic warm eddies (He et al., 2019; Xiu and Chai, 2011).