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Research Article Revised [version 2; peer review: 2 approved, 1 approved with reservations] Samuel J. Toll https://orcid.org/0000-0003-0323-9589 1*, Fiona Qiu https://orcid.org/0000-0003-0642-1116 1*, Yifan Huang https://orcid.org/0000-0003-1060-3668 1, [...] Mark D. Habgood1, Katarzyna M. Dziegielewska https://orcid.org/0000-0002-0070-904X 1, Shuai Nie2, https://orcid.org/0000-0001-6660-7639 1 Samuel J.
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