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Divergence time analysis Because Bayesian dating analysis using BEAST [28] showed little evidence of converging even after a series of multi-month long runs (see below), we applied penalized likelihood in r8s v.1.81 [29] to estimate a chronogram. We calibrated the phylogeny with 28 fossil constraints (Table 1), including the 14 calibration points from Schenk et al. [1]. Criteria for the confidence intervals for the 14 new calibrations can be found in Table 1.
Why some clades are more species-rich than others is a central question in macroevolution. Most hypotheses explaining exceptionally diverse clades involve the emergence of an ecological opportunity caused by a major biogeographic transition or evolution of a key innovation.
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