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1 INTRODUCTION The origin of eusociality is considered one of the major transitions in evolution (Szathmáry & Smith, 1995). Insect societies show a high level of complexity; their colonies have been described as superorganisms (Boomsma & Gawne, 2018; Wheeler, 1911), in which the queen functions as the germline and the workers represent the soma. In ant colonies, for example, queens and workers are closely adapted to their specific roles.
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