1 INTRODUCTION Southwestern China has an extremely unique and complex topographic structure with high species richness. He and Jiang (2014) detailed the phylogeographic structure of sky islands in southwestern China, revealing the factors influencing species evolution and the general pattern of allopatric speciation. The number of species in China has markedly increased in recent years, mainly because of the discovery of new species (Jiang et al., 2015; Wilson & Reeder, 2005).