1 INTRODUCTION The past two decades have witnessed a profound transformation in how morphologists, evolutionary biologists and comparative anatomists study organismal form (Blackburn et al., 2024; Boyer et al., 2016; Davies et al., 2017; Gignac et al., 2024). High-resolution computed tomography (CT), magnetic resonance imaging (MRI) and surface scanning technologies have made it possible to non-destructively capture three-dimensional anatomical structures at resolutions previously unattainable.