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Introduction The Southern Ocean (SO) is an important region globally for ocean circulation and climate (Rintoul and Naveira Garabato, 2013). The Antarctic Circumpolar Current, driven by the Southern Hemisphere westerly winds, forms a zonal flow around the SO, separated by fronts (Sokolov and Rintoul, 2009). The zonal circulation of the SO has long been thought to control the planktonic and benthic biological ecosystems (Linse et al., 2006).
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