aNew York Cooperative Fish and Wildlife Research Unit, Department of Natural Resources, Cornell University, Ithaca, New York, USA. bFisheries, Aquatic Science, and Technology (FAST) Laboratory at Alaska Pacific University, Anchorage, Alaska, USA. dUS Geological Survey, New York Cooperative Fish and Wildlife Research Unit, Cornell University, Ithaca, New York, USA. Received June 14, 2018. Accepted December 18, 2018. Chuenpagdee R, Morgan LE, Maxwell SM, Norse EA, Pauly D. 2003.