Research Articles, Behavioral/Cognitive Tanya Wen, John Duncan and Daniel J Mitchell Journal of Neuroscience 30 September 2020, 40 (40) 7724-7738; DOI: https://doi.org/10.1523/JNEUROSCI.0594-20.2020 Abstract Task episodes consist of sequences of steps that are performed to achieve a goal. We used fMRI to examine neural representation of task identity, component items, and sequential position, focusing on two major cortical systems—the multiple-demand (MD) and default mode networks (DMN).