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1 INTRODUCTION Bumblebees are important pollinators, particularly in temperate regions of the northern hemisphere. However, declines in species abundance and distribution have been reported globally in recent decades (Cameron et al., 2011; Cameron & Sadd, 2020; Williams et al., 2009). According to the IUCN Red List, nearly one-quarter of the bumblebee species that have been assessed are declining (De Meulemeester et al., 2016).
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