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INTRODUCTION Global environmental changes are intensifying, with rising temperatures, frequent flooding, and extreme weather events increasingly threatening biodiversity and global food security (Mace et al., 2008; Urban, 2015; Malanoski et al., 2024). Natural populations typically respond to shifting environments through migration, phenotypic plasticity, selection on standing genetic variation, or de novo mutations (Aitken et al., 2008; Fior et al., 2025).
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