Research Article This article has been accepted for publication and undergone full peer review but has not been through the copyediting, typesetting, pagination and proofreading process, which may lead to differences between this version and the Version of Record. Please cite this article as doi: 10.1002/ps.7630. ABSTRACT BACKGROUND Weeds are among the most damaging pests of agriculture, causing an approximate 10% worldwide reduction in crop productivity each year.