Open AccessArticle by Luke Woodford 1,*, Craig R. Christie 2, Ewan M. Campbell 2, Giles E. Budge 3, Alan S. Bowman 2 and David J. Evans 1 1 Biomedical Sciences Research Complex, University of St. Andrews, St. Andrews KY16 9ST, UK 2 Institute of Biological and Environmental Sciences, School of Biological Sciences, University of Aberdeen, Aberdeen AB24 3FX, UK 3 School of Natural and Environmental Sciences, Newcastle University, Newcastle NE1 7RU, UK * Author to whom correspondence should be addressed.