INTRODUCTION Aphids are an economically important insect group with over 450 species impacting major cropping systems worldwide (Blackman & Eastop, 2017). Over 100 species of the world's aphid fauna are of significant economic importance, transmitting 50% of all insect-borne plant viruses, and they are drawn largely from one family, the Aphididae (Blackman & Eastop 2017; Katis et al., 2007; Nault, 1997).