Hee Ju Yoo

INTRODUCTION Fleshy fruits are crucial for human nutrition and attract frugivores for seed dispersal, with the shift from unripe to ripe fruit representing a key survival strategy for plants (Karlova et al., 2014). Fruit ripening is a highly coordinated process involving physiological and biochemical changes including color, flavor, taste and firmness (Klee & Giovannoni, 2011).