Paolo Moccetti conceived and designed the experiments, performed the experiments, analyzed the data, prepared figures and/or tables, authored or reviewed drafts of the article, and approved the final draft. Jessica R. Rodger conceived and designed the experiments, performed the experiments, authored or reviewed drafts of the article, and approved the final draft. Jonathan D. Bolland conceived and designed the experiments, authored or reviewed drafts of the article, and approved the final draft.