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Abstract Muscle contraction is governed by tropomyosin (Tpm) shifting azimuthally between three states on F-actin (Blocked, Closed and Open) in response to calcium-binding to troponin and acto-myosin cross-bridge formation. The Tpm coiled coil polymerizes head-to-tail along the long-pitch helix of F-actin to form continuous super-helical cables that wrap around the actin filaments.
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