1 INTRODUCTION Niche partitioning can promote the co-occurrence of species in diverse assemblages (Dalsgaard et al., 2013; Dyer et al., 2007; Fründ et al., 2013; Olesen & Jordano, 2002; Schoener, 1974). In assemblages where there is limited niche overlap among species the negative effects of competition on co-occurring species are relaxed (Hutchinson, 1959; Macarthur & Levins, 1967).