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Were early pterosaurs inept terrestrial locomotors?

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By Clark JM, Myers TS, Hsieh ST, Dudek DM
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Introduction The terrestrial competency of pterosaurs was keenly debated during the 1980s and 1990s, when the utility of bipedal and quadrupedal stances, orientation and posture of the extremities, as well as overall terrestriality were discussed at length (Fig. 1, Padian, 1983a; Padian, 1983b; Padian & Olsen, 1984; Wellnhofer, 1988; Unwin, 1988; Unwin, 1989; Unwin, 1996a; Unwin, 1999; Lockley et al., 1995; Bennett, 1997a; Clark et al., 1998; see Witton, 2013 for a recent overview).
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